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The Chronic Myeloproliferative Disorders

The chronic myeloproliferative disorders are characterised by over-production of one or more myeloid cell-lineages. They result from the proliferation of a neoplastic clone of cells derived from a multipotent haemopoietic stem cell, and the leukaemic clone preserves the ability to differentiate and mature. The myeloproliferative disorders differ from the myelodysplastic syndromes, which are largely characterised by quantitative and functional deficiencies in cell production.

Chronic Myeloid Leukaemia

Classification

Chronic Myeloid Leukaemia (CML) is a malignant disorder of the haemopoietic stem cell usually associated with the presence of the Philadelphia chromosome (Ph), a shortened chromosome 22 resulting from a t(9;22) translocation, and at a molecular level by BCR-ABL gene rearrangement. The emergence of the BCR-ABL fusion protein in the stem cell results in the generation of clonal progeny with a selective growth advantage. CML has three recognisable phases; chronic, accelerated, and blastic transformation in which the condition resembles acute leukaemia.

Juvenile CML is a separate clinical condition, not usually associated with the Ph chromosome, and may have trisomy 8, monosomy 7, or be cytogenetically normal. The condition is associated with reversion to foetal haemopoiesis, and does not usually transform to AML.

Clinical Features

Laboratory Diagnosis

Chronic phase: Peripheral blood leucocytosis, reflected in a numerical increase in all stages of myeloid differentiation. Basophilia frequently pronounced. Bone marrow aspirates usually shows profound hypercellularity and increased myeloid:erythroid ratio (>25:1). Philadelphia chromosome present in all myeloid cells.

CML chronic phase peripheral blood; click to enlarge (34K) CML chronic phase bone marrow; click to enlarge (39K)
CML chronic phase: peripheral blood smear
CML chronic phase: bone marrow aspirate
CML chronic phase: bone marrow trephine H & E stain (right). Normal reticulin (left).

Accelerated and blastic phases: Characterised by the emergence of a discrete blast cell population and left-shift compared to chronic phase. Definition of accelerated phase requires at least one of the following:

Blastic phase is defined as >30% blasts and promyelocytes in peripheral blood or marrow. Normal marrow erythropoiesis and megakaryopoiesis reduced.

CML blast transformation: myeloid, with increased basophil precursors; peripheral blood
CML blast transformation: lymphoblastic; peripheral blood
CML blast transformation: megakaryoblastic; peripheral blood, one micro-megakaryoblast
CML megakaryoblastic transformation bone marrow; click to enlarge (39K) CML megakaryoblastic transformation bone marrow; click to enlarge (46K)
CML blast transformation: megakaryoblastic; bone marrow trephine
CML blast transformation: megakaryoblastic with new bone formation (Giemsa stain, right). Bone marrow reticulin stain (left)

Morphology and Cytochemistry

The blast cells seen in accelerated and blastic phases of CML do not fit the established FAB criteria. About 70% of cases in blast transformation have a myeloblast phenotype (CD13, CD33, sometimes erythroid or megakaryocyte markers), 20% have a lymphoblast (Tdt, CD10, CD19, CD22), and 10% a mixed phenotype.

Cytogenetics

Demonstration of the Philadelphia chromosome is performed using standard cytogenetic techniques on leukaemic cells in metaphase. The frequency of the Ph chromosome in marrow samples reflects the degree of marrow replacement by the leukaemic clone. FISH techniques can be used to confirm the presence of BCR-ABL on the Ph chromosome 22 (or rarely chromosome 9), and is particularly useful in Ph- cases of CML, and in cases with complex translocations. Southern- and Western-blot analyses can also be used to demonstrate BCR-ABL. RT-PCR is especially useful in the detection of BCR-ABL for analysis of minimal residual disease.

Prognosis and Treatment

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Juvenile CML

Usually aged 5 or less, predominantly male. The disease is not associated with the Philadelphia chromosome, and produces lower peripheral leucocyte counts than seen in Ph+ CML. There is usually monocytosis and almost always thrombocytopenia. FAB classification of CMML is sometimes recorded. High levels of foetal haemoglobin (Hb-F) are common. 80% of cases lack identifiable genetic abnormalities. Cytotoxic drug treatment is not normally successful, although the disease can be cured by allogeneic, or matched unrelated donor, marrow transplantation.

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Polycythaemia Rubra Vera (PRV)

PRV is an acquired stem cell myeloproliferative disorder, characterised by bone marrow erythroid hyperplasia and increased peripheral blood red-cell volume. Excessive proliferation of the granulocyte and megakaryocyte lines also occurs. PRV has an incidence of 1.8 per 100 000, with an equal male to female ratio.

Symptoms arise from hyperviscocity and vascular damage due to increased red-cell volume. Venous and arterial thromboses are common presenting features, with splenomegaly and hepatomegaly also common.

Laboratory findings usually include raised haemoglobin and packed red-cell volume (haematocrit), but these alone do not constitute PRV. Diagnosis requires the demonstration of a circulating red-cell mass in excess of 36ml/kg, normal arterial O2 saturation, and either splenomegaly, or thrombocytosis (>400 x 109/l) AND leucocytosis (>12 x 109/l). Neutrophils may be 'left-shifted', and absolute basophilia is common. Bone marrow shows erythroid and megakaryocytic hyperplasia, with reduced myeloid:erythroid ratio (<2:1).

PRV; giant platelet in peripheral blood
PRV; hypercellular bone marrow trephine

Median survival in PRV is 9 - 13 years. Treatment includes phlebotomy to reduce the circulating red-cell mass, and hydroxyurea to suppress marrow hyperplasia. 32P and alkylating agents are rarely used now as there is an increased incidence of transformation to acute leukaemia following this type of treatment.

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Myelofibrosis

Myelofibrosis consists of several closely related disorders having several features in common, including progressive marrow fibrosis and extra-medullary haemopoiesis. It results from a proliferation of the fibrous connective tissue which supports haemopoietic elements in the bone marrow. It is believed that the fibroblasts are not abnormal, but may be responding to increased growth factors released from abnormal megakaryocytes.

Myelofibrosis occurs at a frequency about 0.5 per 100 000, and is more common in males than females. Peak age at diagnosis is 60 - 70 years. Splenomegaly is common at presentation, sometimes extensive, resulting from splenic haemopoiesis.

Laboratory diagnosis involves finding evidence of extramedullary haemopoiesis including circulating immature myeloid and erythroid cells, tear-drop poikilocytes (abnormal shaped red-blood cells), and increased bone marrow reticulin fibres without known cause. Giant and bizarrely shaped platelet forms may also occur. Mild leucocytosis with a 'left-shifted' granulocyte series, and mild anaemia due to ineffective erythropoiesis are common at presentation.

Myelofibrosis; peripheral blood showing tear-drop poikilocytes, basophilic stippling and a myelocyte
Myelofibrosis; bone marrow trephine H & E stain (left). Reticulin stain (right)
Osteomyelosclerosis bone marrow; click to enlarge (34K) Osteomyelosclerosis bone marrow; click to enlarge (41K)
Osteomyelosclerosis bone marrow trephine
Osteomyelosclerosis bone marrow reticulin stain

Median survival is about 5 years, although it is likely the disease has been progressing for some years before diagnosis. Poor prognostic factors include severe anaemia, thrombocytopenia and the presence of >10% circulating immature myeloid cells. Mortality is usually due to haemorrhagic or thrombotic complications, and transformation to acute leukaemia occurs in about 20% of cases.

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Essential Thrombocythaemia (ET)

ET is characterised by grossly increased peripheral platelet counts, often exceeding 1 x 1012/l. Diagnosis involves the exclusion of other reactive or malignant conditions associated with very high platelet counts, such as a secondary response to chronic blood loss, chronic inflammatory disorders, and other myeloproliferative disorders. Symptoms results from disturbances in platelet function, and include thromboses and haemorrhage. Moderate splenomegaly is also common.

Laboratory findings include dysmegakaryopoiesis, with giant and bizarrely shaped platelets in the peripheral blood. In contrast to other myeloproliferative disorders, basophilia is not usually present. Bone marrow samples are required to demonstrate marked megakaryocyte hyperplasia, and to investigate the presence of the Ph chromosome, fibrosis, and iron deposits for differentiation from other myeloproliferative disorders.

Essential thrombocythaemia; peripheral blood showing platelet anisocytosis
Essential thrombocythaemia; bone marrow trephine showing clustered megakaryocytes

ET is often an indolent condition, with a long median survival of 12 to 15 years. As ET normally occurs in later years, life expectancy is nearly normal at diagnosis. Treatment normally involves managing the sporadic thromboembolic or haemorrhagic episodes. Transition to acute leukaemia occurs less frequently than in other myeloproliferative disorders.

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Document last updated: Tuesday, 18 November 2003

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